According to Herodotus, the instigator was Aristagoras, tyrant of Miletus, who was trying to cover himself after a failed annexation of the island of Naxos.
Shipwrecked off Mount Athos in the northern Aegean.
The island of Thasos, and Macedon.
Most of the mainland states and all the islands did. Athens and Sparta both forcefully refused.
Earth and water.
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